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4 Xavier Bonnet, Alain Fizesan, Catherine Louise Michel 5 6 7 1. CEBC-CNRS, 79360 Villers en Bois, France 8 9 Correspondance : Xavier Bonnet ; Tel +33 549 097 879 ; Fax +33 549 096 111 ; email 10 [email protected] 11 12 13 Submitted for consideration in Journal of Experimental Biology 14 15 16